Carex

Carex is a vast genus of more than 2,000 species[3] of grass-like plants in the family Cyperaceae, commonly known as sedges (or seg, in older books). Other members of the family Cyperaceae are also called sedges, however those of genus Carex may be called true sedges, and it is the most species-rich genus in the family. The study of Carex is known as caricology.

All species of Carex are perennial,[4] although some species, such as C. bebbii and C. viridula can fruit in their first year of growth, and may not survive longer.[5] They typically have rhizomes, stolons or short rootstocks, but some species grow in tufts (caespitose).[4] The culm – the flower-bearing stalk – is unbranched and usually erect.[4] It is usually distinctly triangular in section.[4]

The leaves of Carex comprise a blade, which extends away from the stalk, and a sheath, which encloses part of the stalk.[4] The blade is normally long and flat, but may be folded, inrolled, channelled or absent.[4] The leaves have parallel veins and a distinct midrib. Where the blade meets the culm there is a structure called the ligule.[4] The colour of foliage may be green, red or brown, and “ranges from fine and hair-like, sometimes with curled tips, to quite broad with a noticeable midrib and sometimes razor sharp edges”.[6]

In this Carex panicea, the upper spike contains male flowers, and the lower spike contains female flowers.
The flowers of Carex are small and are combined into spikes, which are themselves combined into a larger inflorescence. The spike typically contains many flowers, but can hold as few as one in some species.[4] Almost all Carex species are monoecious; each flower is either male (staminate) or female (pistillate).[4] A few species are dioecious. Sedges exhibit diverse arrangements of male and female flowers. Often, the lower spikes are entirely pistillate and upper spikes staminate, with one or more spikes in between having pistillate flowers near the base and staminate flowers near the tip.[7] In other species, all spikes are similar. In that case, they may have male flowers above and female flowers below (androgynous) or female flowers above and male flowers below (gynecandrous). In relatively few species, the arrangement of flowers is irregular.

The defining structure of the genus Carex is the bottle-shaped bract surrounding each female flower.[7] This structure is called the perigynium or utricle, a modified prophyll. It is typically extended into a “rostrum” or beak, which is often divided at the tip (bifid) into two teeth.[7] The shape, venation, and vestiture (hairs) of the perigynium are important structures for distinguishing Carex species.

The fruit of Carex is a dry, one-seeded indehiscent achene or nut[4] which grows within the perigynium. Perigynium features aid in fruit dispersal.